R. van den Bosch

This volume is a revision of Biological Control by R. van den Bosch and P. S. Messenger, originally published by Intext Publishers. In the revision, I have attempted to keep the original theme, and to update it with current research findings and new chapters or sections on insect pathology, microbial control of weeds and plant pathogens, population dynamics, integrated pest management, and economics. The book was written as an undergraduate text, and not as a complete review of the subject area. Various more comprehen­ sive volumes have been written to serve as handbooks for the experts. This book is designed to provide a concise overview of the complex and valuable field of biological control and to show the relationships to the developing concepts of integrated pest management. Population regulation of pests by natural enemies is the major theme of the book, but other biological methods of pest control are also discussed. The chapter on population dynamics assumes a precalculus-levelknowledge of mathematics. Author names of species are listed only once in the text, but all are listed in the Appendix. Any errors or omissions in this volume are my sole responsibility. A. P. Gutierrez Professor of Entomology Division of Biological Control University of California, Berkeley vii Acknowledgments Very special thanks must be given to my colleagues, Professors C. B. Huffaker and L. E. Caltagirone, for the very thorough review they provided and for the many positive suggestions they gave. Dr.

Integrated control of pests was practiced early in this century, well before anyone thought to call it "integrated control" or, still later, "integrated pest management" (IPM), which is the subject of this book by Mary Louise Flint and the late Robert van den Bosch. USDA entomologists W. D. Hunter and B. R. Coad recommended the same principles in 1923, for example, for the control of boll weevil on cotton in the United States. In that program, selected pest-tolerant varieties of cotton and residue destruction were the primary means of control, with insecticides consid­ ered supplementary and to be used only when a measured incidence of weevil damage occurred. Likewise, plant pathologists had also developed disease management programs incorporating varietal selection and cul­ tural procedures, along with minimal use of the early fungicides, such as Bordeaux mixture. These and other methods were practiced well before modern chemical control technology had developed. Use of chemical pesticides expanded greatly in this century, at first slowly and then, following the launching of DDT as a broadly successful insecticide, with rapidly increasing momentum. In 1979, the President's Council on Environmental Quality reported that production of synthetic organic pesticides had increased from less than half a million pounds in 1951 to about 1.4 billion pounds-or about 3000 times as much-in 1977.

This volume is a revision of Biological Control by R. van den Bosch and P. S. Messenger, originally published by Intext Publishers. In the revision, I have attempted to keep the original theme, and to update it with current research findings and new chapters or sections on insect pathology, microbial control of weeds and plant pathogens, population dynamics, integrated pest management, and economics. The book was written as an undergraduate text, and not as a complete review of the subject area. Various more comprehen­ sive volumes have been written to serve as handbooks for the experts. This book is designed to provide a concise overview of the complex and valuable field of biological control and to show the relationships to the developing concepts of integrated pest management. Population regulation of pests by natural enemies is the major theme of the book, but other biological methods of pest control are also discussed. The chapter on population dynamics assumes a precalculus-levelknowledge of mathematics. Author names of species are listed only once in the text, but all are listed in the Appendix. Any errors or omissions in this volume are my sole responsibility. A. P. Gutierrez Professor of Entomology Division of Biological Control University of California, Berkeley vii Acknowledgments Very special thanks must be given to my colleagues, Professors C. B. Huffaker and L. E. Caltagirone, for the very thorough review they provided and for the many positive suggestions they gave. Dr.